What are the Borderlands 4 bosses? Youll find plenty of bosses around Kairos, encountering them in most of the games story missions. The Timekeepers minions consist of a group called The Order, which act as the games main story bosses. While some of the mini-bosses can be more challenging, bosses that are introduced by fancy title cards are the ones youll need to beat to progress through the story.Weve put together a list of all bosses in Borderlands 4, their location, and some tips on how to beat them. Hopefully, youll find some of the Borderlands 4 legendary weapons as you defeat them, and you can check out our guide on the best Borderlands 4 class if youre struggling with any of the fights in the sequel to one of the best FPS games.Read the full story on PCGamesN: All Borderlands 4 bosses and how to beat them